Adam Keith Huttenlocker

Associate Professor of Clinical Integrative Anatomical Sciences

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Dr. Huttenlocker is an Associate Professor in the Department of Integrative Anatomical Sciences where he directs graduate courses in Microscopic Anatomy (IAS 511A & 511B) and serves as the Microanatomy discipline coordinator for Years I & II medical students. He received his PhD from the University of Washington in 2013 and held a National Science Foundation-funded postdoctoral fellowship in comparative vertebrate physiology at The University of Utah from 2013 to 2016.

The Huttenlocker lab at USC investigates the origins of mammalian skeletal function, and applies hard-tissue histology & biomedical imaging techniques to modern and fossil bone in order to interrogate the complex origins of mammalian behavior, growth, and endothermic physiology (warm-bloodedness). The lab also utilizes polarized light microscopy techniques to study collagen fiber orientation in bone and its relevance to bone behavior & function. The lab conducts regular paleontological fieldwork in the southwestern United States (funded by NSF and the Department of the Interior) and southern Africa (NSF and South Africa’s National Research Foundation). Dr. Huttenlocker’s teaching philosophy emphasizes shared patterns in vertebrate development & evolution that explain how & why the body functions the way it does.

Lab Website:


  • Cranial anatomy of Libognathus sheddi Small, 1997 (Parareptilia, Procolophonidae) from the Upper Triassic Dockum Group of West Texas, USA Anat Rec (Hoboken). 2024 Apr; 307(4):1421-1441. . View in PubMed
  • A Carboniferous synapsid with caniniform teeth and a reappraisal of mandibular size-shape heterodonty in the origin of mammals R Soc Open Sci. 2021 Dec; 8(12):211237. . View in PubMed
  • Osteology of the Late Triassic Bipedal Archosaur Poposaurus gracilis (Archosauria: Pseudosuchia) from Western North America Anat Rec (Hoboken). 2020 04; 303(4):874-917. . View in PubMed
  • Vertebrate palaeophysiology Philos Trans R Soc Lond B Biol Sci. 2020 03 02; 375(1793):20190130. . View in PubMed
  • Bone histology of varanopids (Synapsida) from Richards Spur, Oklahoma, sheds light on growth patterns and lifestyle in early terrestrial colonizers Philos Trans R Soc Lond B Biol Sci. 2020 03 02; 375(1793):20190142. . View in PubMed
  • Investigation of a bone lesion in a gorgonopsian (Synapsida) from the Permian of Zambia and periosteal reactions in fossil non-mammalian tetrapods Philos Trans R Soc Lond B Biol Sci. 2020 03 02; 375(1793):20190144. . View in PubMed
  • Histovariability in human clavicular cortical bone microstructure and its mechanical implications J Anat. 2019 11; 235(5):873-882. . View in PubMed
  • Carboniferous-Permian climate change constrained early land vertebrate radiations Nat Ecol Evol. 2019 02; 3(2):200-206. . View in PubMed
  • Publisher Correction: Late-surviving stem mammal links the lowermost Cretaceous of North America and Gondwana Nature. 2018 10; 562(7728):E27. . View in PubMed
  • Late-surviving stem mammal links the lowermost Cretaceous of North America and Gondwana Nature. 2018 06; 558(7708):108-112. . View in PubMed
  • Bone Microvasculature Tracks Red Blood Cell Size Diminution in Triassic Mammal and Dinosaur Forerunners Curr Biol. 2017 Jan 09; 27(1):48-54. . View in PubMed
  • Predator-prey interactions amongst Permo-Triassic terrestrial vertebrates as a deterministic factor influencing faunal collapse and turnover J Evol Biol. 2017 01; 30(1):40-54. . View in PubMed
  • Stem caecilian from the Triassic of Colorado sheds light on the origins of Lissamphibia Proc Natl Acad Sci U S A. 2017 07 03; 114(27):E5389-E5395. . View in PubMed
  • New whaitsioids (Therapsida: Therocephalia) from the Teekloof Formation of South Africa and therocephalian diversity during the end-Guadalupian extinction PeerJ. 2017; 5:e3868. . View in PubMed
  • Breeding Young as a Survival Strategy during Earth’s Greatest Mass Extinction Sci Rep. 2016 Apr 05; 6:24053. . View in PubMed
  • Body size reductions in nonmammalian eutheriodont therapsids (Synapsida) during the end-Permian mass extinction PLoS One. 2014; 9(2):e87553. . View in PubMed
  • An exceptionally preserved transitional lungfish from the lower permian of Nebraska, USA, and the origin of modern lungfishes PLoS One. 2014; 9(9):e108542. . View in PubMed
  • Bone microstructure and the evolution of growth patterns in Permo-Triassic therocephalians (Amniota, Therapsida) of South Africa PeerJ. 2014; 2:e325. . View in PubMed
  • Provincialization of terrestrial faunas following the end-Permian mass extinction Proc Natl Acad Sci U S A. 2013 May 14; 110(20):8129-33. . View in PubMed
  • Comparative anatomy and osteohistology of hyperelongate neural spines in the sphenacodontids Sphenacodon and Dimetrodon (Amniota: Synapsida) J Morphol. 2010 Dec; 271(12):1407-21. . View in PubMed