{"id":43,"date":"2023-08-16T17:16:33","date_gmt":"2023-08-16T17:16:33","guid":{"rendered":"https:\/\/keck.usc.edu\/pibbs\/?page_id=43"},"modified":"2026-09-24T15:51:29","modified_gmt":"2026-09-24T15:51:29","slug":"phd-programs","status":"publish","type":"page","link":"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/","title":{"rendered":"PhD Programs"},"content":{"rendered":"\n\n\n\n\n  \n\n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--hero-primary horizontal-navigation\"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--hero-primary\"\n    \n      >\n\n    \n  \n  <div class=\"inner-wrapper has-horizontal-nav\">\n\n          <div class=\"title-wrapper\">\n\n                  <div class=\"eyebrow-container\">\n                              \n<div class=\"f--field f--eyebrow\">\n\n    \n  <a href=\"https:\/\/keck.usc.edu\/pibbs\/\" >PIBBS Home<\/a>\n\n\n\n<\/div>\n                      <\/div>\n        \n                      \n<div class=\"f--field f--page-title has-stripe\">\n\n    \n      <h1>PhD Programs<\/h1>\n\n\n<\/div>\n        \n              <\/div>\n    \n              \n<div class=\"f--field f--link eyebrow-link\">\n\n    \n    \n  \n\n<a\n  class=\"link \"\n  href=\"https:\/\/keck.usc.edu\/pibbs\/prospective-students\/apply-to-pibbs\/\"\n    >\n        Apply Now to PIBBS\n    <\/a>\n\n\n<\/div>\n    \n    \n      \n              <div class=\"horizontal-nav\">\n                      <div class=\"horizontal-menu-container\">\n              <button class=\"expand-menu\">\n                                  In This Section\n                                <svg class=\"menu\" version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\"\n                  y=\"0px\" viewBox=\"0 0 35 35\" enable-background=\"new 0 0 35 35\" xml:space=\"preserve\">\n                  <path fill-rule=\"evenodd\" clip-rule=\"evenodd\" fill=\"#000\" d=\"M4,23h3v3H4V23z M4,16h3v3H4V16z M10,23\n                  h21v3H10V23z M10,16h21v3H10V16z M10,9h21v3H10V9z M4,9h3v3H4V9z\"\/>\n                <\/svg>\n\n                <svg class=\"close\" version=\"1.1\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" x=\"0px\" y=\"0px\"\n                  viewBox=\"0 0 35 35\" enable-background=\"new 0 0 35 35\" xml:space=\"preserve\">\n                  <polygon fill=\"#000\" fill-rule=\"evenodd\" clip-rule=\"evenodd\" points=\"26,10.4 24.6,9 17.5,16.1 10.4,9 9,10.4 16.1,17.5\n                  9,24.6 10.4,26 17.5,18.9 24.6,26 26,24.6 18.9,17.5 \"\/>\n                <\/svg>\n              <\/button>\n\n              <nav\n  class=\"mc--menu mc--\"\n  >\n  <ul class=\"m--menu m--\">\n    \n    \n      <li class=\"has-submenu menu-item menu-item-type-post_type menu-item-object-page menu-item-home menu-item-7 menu-item-has-children\">\n      <div class=\"link-arrow-wrapper\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/\" >PIBBS<\/a>\n\n      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menu-item-type-post_type menu-item-object-page menu-item-36\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/prospective-students\/apply-to-pibbs\/\" >Apply to PIBBS<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-40\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/prospective-students\/recruitment-interview-visit\/\" >Recruitment Visit<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-42\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/recruitment-schedule\/\" >Recruitment Schedule<\/a>\n  \n  \n<\/li>\n                  <\/ul>\n      <\/div>\n    <\/div>\n  \n<\/li>\n\n    \n      <li class=\"has-submenu menu-item menu-item-type-post_type menu-item-object-page menu-item-44 menu-item-has-children\">\n      <div class=\"link-arrow-wrapper\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/\" >PhD Programs<\/a>\n\n      <button class=\"expand-children has-submenu arrow-toggle\" aria-expanded=\"false\" aria-haspopup=\"true\" aria-label=\"Display Submenu for PhD Programs\">\n        <svg height=\"35\" viewBox=\"0 0 35 35\" width=\"35\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\">\n          <path d=\"m17.5 20.2142857 6.2142857-6.2142857 1.2857143 1.2857143-7.5 7.5-7.5-7.5 1.2857143-1.2857143z\" fill=\"#000\" fill-rule=\"evenodd\"\/>\n        <\/svg>\n      <\/button>\n    <\/div>\n  \n      <div class=\"submenus-wrapper\">\n      <div class=\"submenus-wrapper-inner\">\n        <ul class=\"submenu\" aria-hidden=\"true\">\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-46\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/cancer-biology-and-genomics\/\" >Cancer Biology and Genomics<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-54\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/development-stem-cell-and-regenerative-medicine\/\" >Development, Stem Cells and Regenerative Medicine<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-61\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/infectious-diseases-immunology-pathogenesis\/\" >Infectious Diseases, Immunology and Pathogenesis<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-71\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/medical-biophysics\/\" >Medical Biophysics<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-195\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/molecular-medicine\/\" >Molecular Medicine<\/a>\n  \n  \n<\/li>\n                  <\/ul>\n      <\/div>\n    <\/div>\n  \n<\/li>\n\n    \n      <li class=\"has-submenu menu-item menu-item-type-post_type menu-item-object-page menu-item-258 menu-item-has-children\">\n      <div class=\"link-arrow-wrapper\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/pibbs-faculty\/\" >PIBBS Faculty<\/a>\n\n      <button class=\"expand-children has-submenu arrow-toggle\" aria-expanded=\"false\" aria-haspopup=\"true\" aria-label=\"Display Submenu for PIBBS Faculty\">\n        <svg height=\"35\" viewBox=\"0 0 35 35\" width=\"35\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\">\n          <path d=\"m17.5 20.2142857 6.2142857-6.2142857 1.2857143 1.2857143-7.5 7.5-7.5-7.5 1.2857143-1.2857143z\" fill=\"#000\" fill-rule=\"evenodd\"\/>\n        <\/svg>\n      <\/button>\n    <\/div>\n  \n      <div class=\"submenus-wrapper\">\n      <div class=\"submenus-wrapper-inner\">\n        <ul class=\"submenu\" aria-hidden=\"true\">\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-272\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/pibbs-faculty\/cbg-faculty\/\" >CBG Faculty<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-271\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/pibbs-faculty\/dsr-faculty\/\" >DSR Faculty<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-269\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/pibbs-faculty\/idip-faculty\/\" >IDIP Faculty<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-270\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/pibbs-faculty\/mbph-faculty\/\" >MBPH Faculty<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-277\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/pibbs-faculty\/mmed-faculty\/\" >MMED Faculty<\/a>\n  \n  \n<\/li>\n                  <\/ul>\n      <\/div>\n    <\/div>\n  \n<\/li>\n\n    \n      <li class=\"has-submenu menu-item menu-item-type-post_type menu-item-object-page menu-item-77 menu-item-has-children\">\n      <div class=\"link-arrow-wrapper\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/student-resources\/\" >Student Resources<\/a>\n\n      <button class=\"expand-children has-submenu arrow-toggle\" aria-expanded=\"false\" aria-haspopup=\"true\" aria-label=\"Display Submenu for Student Resources\">\n        <svg height=\"35\" viewBox=\"0 0 35 35\" width=\"35\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\">\n          <path d=\"m17.5 20.2142857 6.2142857-6.2142857 1.2857143 1.2857143-7.5 7.5-7.5-7.5 1.2857143-1.2857143z\" fill=\"#000\" fill-rule=\"evenodd\"\/>\n        <\/svg>\n      <\/button>\n    <\/div>\n  \n      <div class=\"submenus-wrapper\">\n      <div class=\"submenus-wrapper-inner\">\n        <ul class=\"submenu\" aria-hidden=\"true\">\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-84\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/student-resources\/student-forms\/\" >Student Forms<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-86\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/student-resources\/student-involvement\/\" >Student Involvement<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-89\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/student-resources\/pibbs-alumni\/\" >PIBBS Alumni<\/a>\n  \n  \n<\/li>\n                      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-122\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/student-resources\/pibbs-spotlights\/\" >PIBBS Spotlights<\/a>\n  \n  \n<\/li>\n                  <\/ul>\n      <\/div>\n    <\/div>\n  \n<\/li>\n\n    \n      <li class=\" menu-item menu-item-type-post_type menu-item-object-page menu-item-174\">\n      <a href=\"https:\/\/keck.usc.edu\/pibbs\/contact\/\" >Contact<\/a>\n  \n  \n<\/li>\n\n    \n      <\/ul><!-- m--menu -->\n<\/nav>\n\n            <\/div>\n                  <\/div>\n        <\/div>\n\n\n  <\/div><\/div>\n\n\n\n\n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--rich-text white\"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--rich-text\"\n    \n      >\n\n    \n  <div class=\"inner-wrapper\">\n        \n<div class=\"f--field f--wysiwyg\">\n\n    \n  <h3 class=\"fusion-responsive-typography-calculated\" data-fontsize=\"30\" data-lineheight=\"45px\">PhD Programs in Biomedical and Biological Sciences (PIBBS)<\/h3>\n<p><img loading=\"lazy\" decoding=\"async\" class=\"aligncenter size-medium wp-image-172\" src=\"https:\/\/keck.usc.edu\/pibbs\/wp-content\/uploads\/sites\/27\/2023\/08\/pibbs-programs-300x159.jpg\" alt=\"Portrait of five PIBBS students.\" width=\"300\" height=\"159\" srcset=\"https:\/\/keck.usc.edu\/pibbs\/wp-content\/uploads\/sites\/27\/2023\/08\/pibbs-programs-300x159.jpg 300w, https:\/\/keck.usc.edu\/pibbs\/wp-content\/uploads\/sites\/27\/2023\/08\/pibbs-programs-1024x541.jpg 1024w, https:\/\/keck.usc.edu\/pibbs\/wp-content\/uploads\/sites\/27\/2023\/08\/pibbs-programs-768x406.jpg 768w, https:\/\/keck.usc.edu\/pibbs\/wp-content\/uploads\/sites\/27\/2023\/08\/pibbs-programs.jpg 1500w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/><\/p>\n<p><span style=\"font-weight: 400;\">Our goal is to recruit top caliber PhD students in the biomedical and biological sciences to USC and guide them toward a mentor, lab, and research project. From there, we support each student through rigorous scientific training, an outstanding PhD thesis, and a strong foundation for a variety of science-related careers. Six interdisciplinary PhD programs make up PIBBS.<\/span><\/p>\n\n\n\n<\/div>\n  <\/div>\n\n\n  <\/div><\/div>\n\n\n\n\n\n\n          \n  \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--accordions \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--accordions\"\n    \n      >\n\n    \n      <div class=\"header-container\">\n\n                  \n<div class=\"f--field f--section-title\">\n\n    \n  <h2>\n          Five PhD Tracks\n      <\/h2>\n\n\n<\/div>\n      \n                  \n<div class=\"f--field f--description description-large\">\n\n    \n  <p>For additional information on any track, please visit the PhD&#8217;s site page an see the <a href=\"https:\/\/catalogue.usc.edu\/\" target=\"_blank\" rel=\"noopener\">USC Course Catalog<\/a>.<\/p>\n\n\n\n<\/div>\n      \n    <\/div>\n  \n      <ul>\n              <li>\n          <button type=\"button\" class=\"accordion-trigger \" id=\"heading-1-1-ls6ZWtzaqu\" aria-controls=\"section-1-1-ls6ZWtzaqu\" aria-expanded=\"false\" aria-disabled=\"false\">\n                          <span class=\"item-title\">Cancer Biology &#038; Genomics (CBG)<\/span>\n            \n                      <\/button>\n\n          <div id=\"section-1-1-ls6ZWtzaqu\" role=\"region\" aria-labelledby=\"heading-1-1-ls6ZWtzaqu\" class=\"accordion-panel\">\n\n                            \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--rich-text \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--rich-text\"\n    \n      >\n\n    \n  <div class=\"inner-wrapper\">\n        \n<div class=\"f--field f--wysiwyg\">\n\n    \n  <p><a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/cancer-biology-and-genomics\/\">CBG<\/a> capitalizes on our<a href=\"http:\/\/uscnorriscancer.usc.edu\/\"> Norris Comprehensive Cancer Center<\/a>, one of the oldest National Cancer Institute-designated Comprehensive Cancer Centers in the United States. A Comprehensive Cancer Center designation requires that clinical treatment of patients and scientific research are carried out side by side. The Norris has strength in all areas of cancer research, including clinical and basic research and epidemiology.<\/p>\n\n\n\n<\/div>\n  <\/div>\n\n\n  <\/div><\/div>\n            \n                      <\/div>\n        <\/li>\n\n              <li>\n          <button type=\"button\" class=\"accordion-trigger \" id=\"heading-1-2-ls6ZWtzaqu\" aria-controls=\"section-1-2-ls6ZWtzaqu\" aria-expanded=\"false\" aria-disabled=\"false\">\n                          <span class=\"item-title\">Development, Stem Cells, &#038; Regenerative Medicine (DSR)<\/span>\n            \n                      <\/button>\n\n          <div id=\"section-1-2-ls6ZWtzaqu\" role=\"region\" aria-labelledby=\"heading-1-2-ls6ZWtzaqu\" class=\"accordion-panel\">\n\n                            \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--rich-text \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--rich-text\"\n    \n      >\n\n    \n  <div class=\"inner-wrapper\">\n        \n<div class=\"f--field f--wysiwyg\">\n\n    \n  <p><a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/development-stem-cell-and-regenerative-medicine\/\">DSR<\/a> benefits from our wonderful Center for Regenerative Medicine and our strength in stem cell research, tissue engineering and regeneration.<\/p>\n\n\n\n<\/div>\n  <\/div>\n\n\n  <\/div><\/div>\n            \n                      <\/div>\n        <\/li>\n\n              <li>\n          <button type=\"button\" class=\"accordion-trigger \" id=\"heading-1-3-ls6ZWtzaqu\" aria-controls=\"section-1-3-ls6ZWtzaqu\" aria-expanded=\"false\" aria-disabled=\"false\">\n                          <span class=\"item-title\">Infectious Diseases, Immunology &#038; Pathogenesis (IDIP)<\/span>\n            \n                      <\/button>\n\n          <div id=\"section-1-3-ls6ZWtzaqu\" role=\"region\" aria-labelledby=\"heading-1-3-ls6ZWtzaqu\" class=\"accordion-panel\">\n\n                            \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--rich-text \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--rich-text\"\n    \n      >\n\n    \n  <div class=\"inner-wrapper\">\n        \n<div class=\"f--field f--wysiwyg\">\n\n    \n  <p><a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/infectious-diseases-immunology-pathogenesis\/\">IDIP<\/a> is a program focused on translating bench research to the bedside by investigating human organ systems in health and disease. It takes advantage of our outstanding clinical and basic scientists and the exciting interdisciplinary and collaborative atmosphere at USC.<\/p>\n\n\n\n<\/div>\n  <\/div>\n\n\n  <\/div><\/div>\n            \n                      <\/div>\n        <\/li>\n\n              <li>\n          <button type=\"button\" class=\"accordion-trigger \" id=\"heading-1-4-ls6ZWtzaqu\" aria-controls=\"section-1-4-ls6ZWtzaqu\" aria-expanded=\"false\" aria-disabled=\"false\">\n                          <span class=\"item-title\">Medical Biophysics (MBPH)<\/span>\n            \n                      <\/button>\n\n          <div id=\"section-1-4-ls6ZWtzaqu\" role=\"region\" aria-labelledby=\"heading-1-4-ls6ZWtzaqu\" class=\"accordion-panel\">\n\n                            \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--rich-text \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--rich-text\"\n    \n      >\n\n    \n  <div class=\"inner-wrapper\">\n        \n<div class=\"f--field f--wysiwyg\">\n\n    \n  <p><a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/medical-biophysics\/\">MBPH<\/a> is a program seeks to answer important biomedical questions using all aspects of biophysics, with an emphasis on structural biology and biophysical methods and theories. Some faculty members of this program are located on the University Park (undergraduate) USC campus.<\/p>\n\n\n\n<\/div>\n  <\/div>\n\n\n  <\/div><\/div>\n            \n                      <\/div>\n        <\/li>\n\n              <li>\n          <button type=\"button\" class=\"accordion-trigger \" id=\"heading-1-5-ls6ZWtzaqu\" aria-controls=\"section-1-5-ls6ZWtzaqu\" aria-expanded=\"false\" aria-disabled=\"false\">\n                          <span class=\"item-title\">Molecular Medicine (MMED)<\/span>\n            \n                      <\/button>\n\n          <div id=\"section-1-5-ls6ZWtzaqu\" role=\"region\" aria-labelledby=\"heading-1-5-ls6ZWtzaqu\" class=\"accordion-panel\">\n\n                            \n    \n\n\n\n\n\n\n<div\n  class=\"cc--component-container cc--rich-text \"\n\n  \n  \n  \n  \n  \n  \n  >\n  <div class=\"c--component c--rich-text\"\n    \n      >\n\n    \n  <div class=\"inner-wrapper\">\n        \n<div class=\"f--field f--wysiwyg\">\n\n    \n  <p><a href=\"https:\/\/keck.usc.edu\/pibbs\/phd-programs\/molecular-medicine\/\">MMED<\/a> offers comprehensive research training at the intersection of medicine and molecular biology, equipping students with practical skills and a collaborative environment to make significant contributions to medical advancements.<\/p>\n\n\n\n<\/div>\n  <\/div>\n\n\n  <\/div><\/div>\n            \n                      <\/div>\n        <\/li>\n\n          <\/ul>\n  \n\n  <\/div><\/div>\n","protected":false},"excerpt":{"rendered":"","protected":false},"author":3,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_acf_changed":false,"advgb_blocks_editor_width":"","advgb_blocks_columns_visual_guide":"","footnotes":""},"class_list":["post-43","page","type-page","status-publish","hentry"],"acf":[],"yoast_head":"<!-- This site is optimized with the Yoast SEO Premium plugin v28.2 (Yoast SEO v28.2) - 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